# The FossilSim package

#### Joelle Barido-Sottani, Rachel C. M. Warnock

#### 2025-10-28

This vignette provides a Quick start and a brief introduction to the package input and output.

## Contents

- Background and motivation
- Installing the package
- Quick start
- FossilSim input
- FossilSim output
- What next?

## Background and motivation

`FossilSim` is an R package for simulating species taxonomy and fossil occurrence data in a phylogenetic framework. Simulations can be used to address many questions in phylogenetics and palaeobiology, and are especially useful for assessing the performance of different methods, since the true underlying parameters (e.g. diversification and fossil recovery rates) are known. Tara Smiley provides an excellent demonstration of the value of simulations in palaeobiology in this blog post. `FossilSim` output can be easily parsed for downstream analysis using R, or other software packages including BEAST2 and RevBayes. The package provides a wide range of flexible models, including models of interval-, lineage- and environment-dependent fossil recovery, in addition to plotting functions that can be used to visualise the output and produce publication quality figures.

## Installing the package

`FossilSim` is available on CRAN or the latest development version can be installed from GitHub using the package `devtools`.

    # to install the package via CRAN
    install.packages("FossilSim")

    # to install the package via GitHub
    devtools::install_github("rachelwarnock/fossilsim")

    # load the package into the current working environment
    library(FossilSim)

### Package dependencies

The installation above will automatically install the package dependencies, `ape` and `TreeSim`.

### Calling functions from FossilSim and other packages

Once the package is loaded into your current working environment you can call the package functions directly, e.g. `sim.fossils.poisson()`. It is also possible to call functions in R without loading the package into your working environment, e.g. `FossilSim::sim.fossils.poisson()`. Throughout this vignette and other documentation associated with `FossilSim`, we call the `FossilSim` functions from the current working environment but use the `::` format to call functions from other packages. This is done to make the source of all functions as clear as possible.

### Quick start

Simulating data using `FossilSim` can be as simple as the following code snippets.

    # simulate a tree using ape
    tips = 8
    t = ape::rtree(tips)
    # simulate fossils using fossilsim
    rate = 2
    f = sim.fossils.poisson(rate = rate, tree = t)
    # plot the output
    plot(f, tree = t)

    # simulate taxonomy using fossilsim
    beta = 0.5 # probability of symmetric speciation
    lambda.a = 0.1  # rate of anagenesis
    s = sim.taxonomy(tree = t, beta = beta, lambda.a = lambda.a)
    # plot the output
    plot(s, tree = t, legend.position = "bottomright")

The package contains many options for simulating fossils and evaluating the output in a meaningful way requires understanding the underlying model parameters. Parsing the output for downstream analysis also requires becoming familiar with the `FossilSim` objects.

## FossilSim input

The starting point for any data generation using `FossilSim` is a phylogenetic tree. `FossilSim` relies on the widely used `ape` package `phylo` object format for handling trees. The `phylo` object stores information about the relationship among branches in a phylogeny in a matrix called `edge` and branch lengths are stored in a vector called `edge.length`. The age of the tips and internal nodes can be reconstructed by combining information from the `edge` matrix and the `edge.length` vector. If the tree has a root edge (i.e. a branch leading to the root) the length of this edge is stored as a numeric variable called `root.edge`.

There are a huge number of packages and options for simulating trees that can be used as input in `FossilSim`, including the `ape` and `TreeSim` R packages. An empirical phylogeny can also be used as input. The only general requirements are that trees are fully resolved and scaled to time.

## FossilSim output

`FossilSim` can be used to simulate species taxonomy and fossil sampling times, which are stored and output as the `taxonomy` and `fossils` objects described in the “Simulating taxonomy” and “Simulating fossils” vignettes. The functions used to simulate fossils can take either a `phylo` or `taxonomy` object as input. This means, in theory, the user never has to interact with the `taxonomy` object when simulating fossil data. However, it may still be useful to become familiar with the concepts underlying the `taxonomy` object.

## What next?

Information about the `taxonomy` object and models for simulating taxonomy can be found in the vignette “Simulating taxonomy”. Information about the `fossils` object and models for simulating fossils can be found in the vignette “Simulating fossils”. See the `paleotree` vignette to see how `FossilSim` objects can be converted into `paleotree` objects.